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However, there is little all dispersed out of an area of origin in the An- controversy regarding the existence of clocks des of Bolivia or south eastern Peru, with suc- which work well within specific taxonomic cessive pulses of dispersal northwards and in- groups Li We therefore see no problem termittent isolation of populations during the in using DNA sequences of related organisms last twoglacial periods.

A fairly detailed se- to investigate their history, and in fact several quence of events is suggested. The results and other molecular studies of Andean birds sug- interpretations of a molecular phylogenetic gest that vicariance patterns that are prominent study of the same group Garcfa-Moreno et al. This stu- We would like to comment Furthermore, based on several years DNA hy- on some of the discrepancies between these bridisation work, Bleiweiss a, b sug- two analyses of the same taxonomic group.

Our estimates with these different rates tion. The alternative would be to base the time are similar and both point to a divergence be- estimates on apre-conceived idea about the role tween mid elevation and highland species of Pleistocene glaciations, which does not offer about 2 - 4 mya. This result holds for us even a mechanism for critical hypothesis evaluation.

However, 3 The position of M. The DNA evidence interpretations based on a crude phylogeny and supports its placement in a clade together the current distribution patterns cannot lead be- with geographically adjacent forms living in yond the speculative level of scenario building south-western Ecuador thus suggesting a vi- and can never provide a mechanism for rigorous cariance pattern even if the branching pattern evaluation of ideas e.

The only evidence they produced was to refer to the existence of a melanistic Boli- forkedtail,monornorph phoebe vian species in the genus Aglaeactis. Why theresiae not refer to the existence of a melanistic Coe- ligena in north-western Colombia?

The char- --aeneocauda acters they used to infer a phylogeny can be 67rEcuador mapped without conflict into the DNA gener- ated phylogeny Fig. However, we find it peculiar that they use this to build a Bolivia detailed narrative with a specific time scale, supporting a general idea about a Bolivian dis There is considerable evidence sug- Fig. Maximum likelihood tree obtained from gesting random segregation of pigment charac- bp of combined DNA sequences of mitochon- ters in birds inhabiting patchy Andean habitats drial cyt b bp , ND2 bp and ND5 e.

Figures above the tallura case the authors were aware that an al- branches represent bootstrap values with the maxi- ternative interpretation exists and is supported mum likelihood algorithm after replicates. We earlier proposed Branches are drawn proportionate to the amount of that the Andean avifauna diversified in the change along the branch, and those with low boot- upper Pleistocene see e.

Names as attempts critically to evaluate a model Roy in the M. We do not think Carchi and Loja. Mougin - Pairing in Cow's Shearwater ply fitting phenotypic variation into a precon- implications for the speciation process.

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